Summary Information about The Roslin Institute.

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1 Professor David A Hume Director, The Roslin Institute and Research Director, Royal (Dick) School of Veterinary Studies University of Edinburgh Professor David Argyle Dean, Royal (Dick) School of Veterinary Studies Ms Val White Campus Operating Officer About The Roslin Institute The vision of The Roslin Institute is to be the leading Animal Sciences Institute in the world. The Roslin Institute is a Biotechnology and Biological Sciences Research Council (BBSRC) National Institute of Bioscience incorporated with the University of Edinburgh s Royal (Dick) School of Veterinary Studies (R(D)SVS), the number one-ranked Veterinary School in the UK in the 2008 Research Assessment Exercise. The Roslin Institute undertakes research focussed on the health and welfare of animals, and applications of basic animal sciences in human and veterinary medicine, the livestock industry and food security. In April 2007, Roslin Institute was integrated with the Neuropathogenesis Unit (formerly of the Institute for Animal Health), and in April 2008 the combined organisation affiliated with the R(D)SVS within the College of Medicine and Veterinary Medicine (CMVM) of the University of Edinburgh. In 2011, The Roslin Institute, with the Animal Sciences Division of Scotland s Rural College, occupied a new, state-of-the-art, 60M research building that forms a part of the University of Edinburgh s Easter Bush Veterinary Campus (EBVC), also including the new teaching facility for Veterinary Studies, small and large animal hospitals and the Oncology centre. This places The Roslin Institute alongside the R(D)SVS, and Easter Bush Research Consortium (EBRC) partners Scotland s Rural College and the Moredun Research Institute. The Roslin Institute has over 70 Group Leaders and staff and students affording a globally-leading critical mass to deliver the highest international quality research and postgraduate training in animal biosciences. Its state-of-the-art, cost effective research infrastructure provides access to the latest technologies and skills across a wide range of cutting-edge disciplines. Research activity at The Roslin Institute underpins >1,321 jobs, with every 1 spent is returned into the UK economy, and continues to generate productivity gains in the UK livestock and food production sector (est. 320M GVA since the 1960s). The Roslin Institute s genetic facilities to date have been used by over 1,137 individuals from over 46 countries. The Roslin Institute has a long- established track record of industrial partnerships with over 75% of its staff having some form of industrial link, it currently has active collaborations with over 40 companies; this expertise will be strengthened, developed and extended to enhance the collaborative opportunities available and increase the economic impact of its research accordingly. The Overall Objectives of The Roslin Institute and R(D)SVS The Roslin Institute and R(D)SVS, in cooperation with the EBRC and College of Medicine and Veterinary Medicine (CMVM), aims to improve the lives of animals and humans through world class research in animal biology and to make the one biology, one medicine concept of the College into a reality. 1 Figures as per Biggar Economics Jan 2013

2 The principal research objectives are: Improved animal health and welfare through knowledge of genetic factors affecting susceptibility and resistance to disease Improved sustainability of livestock production systems and food supply chains through an understanding of the biological, economic, environmental and social factors that apply, and their interactions Translation of discoveries into veterinary clinical practice, and translation of veterinary clinical observations and opportunities into basic science discoveries. Improved food safety based upon understanding interactions between disease causing organisms and animals Improved human health through an understanding basic mechanisms of health and disease through comparative biology of animal species the identification of new and emerging zoonoses and an improved understanding of how pathogens might cross from animals to humans Improved quality of life for animals by studying the mechanisms and behaviours associated with optimising their environment and life experiences. BBSRC-Funded Institute Strategic Programme Grants Research at The Roslin Institute contributes is based upon a set of five BBSRC-funded Institute Strategic Programmes: Programme 1: Analysis and Prediction in Complex Animal Systems Aim: To combine leading research in quantitative genetics and animal genomics with the development of tools and resources for science and for knowledge exchange and commercialisation. The key deliverables of this programme are high-quality reference genomes for major livestock species, new ways of interpreting and presenting this information, a firm foundation for livestock expression atlases, novel dissections of genetic variance using systems approaches, improved outcomes from genomic selection, and new ways of using field data to identify genetic variation in disease. Programme 2: Control of Development and Reproduction Traits Aim: To investigate aspects of development, growth and reproduction of farm animal species advancing our understanding of production traits and basic bioscience underpinning animal and human health and wellbeing. This will be achieved using the power of the genetic and genomics resources available in farm animals and informative model species, linking this to functional studies in whole animals and at the cellular and molecular levels. In parallel we will develop and apply novel tools and resources, particularly in stem cells and genetic modification technologies, which are based on The Roslin Institute s established, world-leading expertise. This programme supports, and is supported by, the National Avian Research Facility. As an interdisciplinary programme pursuing targeted basic research, while aiming to meet both industry needs and consumer expectations, this programme addresses the challenges of 'Food Security' and 'Basic Bioscience Underpinning Health', central to the "One Biology, One Health" agenda. Programme 3: Innate Immunity and Endemic Disease Aim: The Roslin Institute is the only UK Institute with a substantial programme on endemic non-viral diseases of livestock. Each major livestock species can be infected with a range of pathogens, each contributing to a substantial economic burden. Many of these pathogens can also infect humans, and are major public health concerns. The global aim of this programme is to reduce the economic burden of endemic non-viral diseases of livestock and to mitigate impacts on food safety. The research aims to carry out research into host-pathogen interactions that can be applied to reduce the threat from infectious diseases to the UK, thereby improving the sustainability of UK agriculture and reducing our dependence on imported food. As many human infections originate from food and production animals, interventions to mitigate these diseases in the host animal population should also reduce the burden of human disease. This

3 programme is based upon fundamental research on the genetics, genomics and cell biology of innate immunity in livestock, and includes research on livestock responses, host susceptibility and vaccine development to important bacterial zoonoses. Programme 4: Pathogenesis and Resistance in Viral Diseases of Livestock Aim: This programme deals with the immunology, genetics and genomics of host resistance and susceptibility to virus infection and disease, outstanding fundamental questions on virus pathogenesis and novel strategies for the control of virus diseases. Our aim is to address important gaps in fundamental knowledge or in specific diseases and to translate the outputs into novel control strategies including animal breeding, transgenics, new vaccines or vaccine strategies. The programme includes detailed studies on a wide variety of mainly endemic viruses of importance to food production in their natural hosts, some of them with significant potential as vaccine vectors. The programme brings together researchers who specialise in a wide range of pathogens and specialists in the immune systems of livestock, and in the genetics, genomics and pathways of disease resistance. The strength of this aggregation is seen in our ability to study the impacts of co-infections, and in comparative pathogenesis, applying the knowledge of ways that pathogens subvert host resistance pathways to understanding the biology of other pathogens. Programme 5: Livestock Neurobiology. Aim: This programme has grown out of groups that study the biology of transmissible spongiform encephalopathies (TSEs) (including BSE and variant CJD). A focus on TSEs remains critical in the face of continuing and new threats from this class of diseases. Additionally, within this programme we will focus on the role of proteins of the prion family and the immune and other glial cells of the CNS in the maintenance of neuronal health, particularly during ageing. We will also investigate the way that the neuroendocrine system allows neurons to respond to diverse and physiologically-relevant types of stress, including social stress; understanding such mechanisms is a prerequisite to developing strategies for reversing neuronal loss. This aspect is at the heart of Basic Bioscience Underpinning Health and addresses community concerns about animal welfare and behaviour that also influence sustainable productivity. Although livestock animals are not aged to the same extent as humans, they experience a range of different challenges during life, including poor diets, social stress and intensive farming. Maintaining animals in a healthy state in the face of such stress is of crucial importance to underpin the agricultural industry. In addition to the five Institute Strategic Programme Grants that underpin The Roslin Institute s research, the Institute is proud to host to two National Capabilities: The ARK-Genomics Centre for Comparative and Functional Genomics ARK-Genomics is the major UK Centre for livestock genomic analysis with capabilities including sequencing, genotyping, transcriptomics and informatics. ARK-Genomics has been operating as an international facility for 10 years and we maintain contact with 1,137 individuals, across 504 companies/departments and 46 countries. As part of ARK Genomic s role as a resource centre, we have material distribution agreements (MDAs) with 170 companies/departments internationally. Of the 193 collaborative research partners outside of The Roslin Institute, 48% are based within the UK, 34% within Europe and 18% outside Europe, proving both national and international need and impact. This facility underpins much of the research of The Roslin Institute. ARK-Genomics uses the latest genomics technologies for research into genome structure, genetic variation, gene expression and gene function with a focus on systems of relevance to animal health and food security. The laboratory employs a range of technologies, including next-generation sequencing, SNP genotyping, microarrays, and bioinformatics, as well as managing and distributing an extensive and unique clone/vector library. ARK-Genomics combines the latest technologies with bioinformatics research to provide a complete genomics solution from concept to conclusion. ARK-Genomics focuses on the quantitative biology, genetics and genomics of food-producing animals, their model systems and their pathogens. This focus covers the BBSRC priority areas of "Food Security" and "Basic Bioscience underpinning Health". The objectives are to

4 investigate the genetic factors that influence yield, animal production, food safety, resistance to pathogens, and the immune response, as well as basic science underpinning the search for drug targets and vaccine development. The outputs feed directly into the breeding and other translational programmes of industrial and academic partners. With a track record in animal sciences and the close integration with The Roslin Institute, ARK-Genomics addresses the challenges of food production and animal disease, and the increasing global demand for food production and security. In 2013 ARK-Genomics and The GenePool at the School of Biological Sciences merged to form Edinburgh Genomics. This merger achieved two objectives. It made the joint facility agile and robust, able to take on projects from the very large to the very small across the gamut of approaches in modern genomics. It also allows proactive investment in the newest technologies and to offer the economies of scale to our collaborators. Together the Edinburgh Genomics teams have been delivering sequence data and analysis to researchers for nearly 20 years, and next generation data since It has a wealth of experience, and a record of delivery, that cements its position as one of the leading genomics facilities in the UK. National Avian Research Facility The UK is at the forefront of research into the biology of the chicken. Central to this, The Roslin Institute s work on poultry is a major component of the Institute s research, and increasingly birds are viewed as model organisms for basic bioscience in addition to their obvious economic importance. The National Avian Research Facility will maintain and provide access to inbred and transgenic lines held under specified pathogen free (SPF) conditions and transgenic and germ cell preservation technologies alongside genetic/genomic analytical resources. The National Avian Research Facility will provide access to the latest cutting-edge technologies and unique lines of chickens that will allow research into the biology of disease in the chicken and the development of novel vaccines and therapeutics that will impact on human health (chickens are reservoirs for human pathogens such as Salmonella and Campylobacter (both food-borne bacterial pathogens that generally cause food poisoning but can have serious consequences) and of course avian influenza) but also on the health and welfare of the approximately one billion chickens raised in the UK every year. They will also allow us to try to understand the role of genes and pathways in embryonic development, for which the chicken is a widely used model. Another potential benefit of the research to be carried out in, and facilitated by, the facility is the ability to cryopreserve different lines of chickens and ultimately different species of birds, particularly those under threat of extinction. Such a "biobank" is now theoretically possible and would preserve species and genetic diversity that would otherwise be lost forever. Research will continue into the relevant technologies in this facility to try to turn theory into practice. The SPF Avian Facility will be jointly managed by The Roslin Institute and The Pirbright Institute (another Institute that receives strategic funding from the BBSRC). Inbred poultry lines currently housed at The Pirbright Institute will be transferred to Easter Bush. The Pirbright Institute is developing a new hatchery and high containment challenge facility. This high containment facility is part of The Pirbright Institute National Capability and will be available for use both by Pirbright and Roslin Institute research groups and those from other UK institutions including the avian infectious diseases groups at The Roslin Institute who will use the Pirbright facilities for their high containment work. Both Roslin and Pirbright Institutes work on virus infections of livestock and poultry but broadly-speaking, while Pirbright works across many aspects of viral diseases, The Roslin Institute comes to viral diseases from the side of the host; working on the impact of genetic variation on virus disease-resistance and the pathways involved in host defence.

5 The Roslin Institute also hosts a number of Core Technology Units including: Small and Large Animal Units Computational Biology and Bioinformatics Proteomics and Mass Spectrometry Protein Expression and Antibody Production Transgenic Animal Technologies Visualisation and Imaging Recent Roslin Institute achievements New partnership with Cobb-Vantress Inc announced A three-year research agreement between Cobb-Vantress, Inc., a global industry leader in poultry genetics, and The Roslin Institute, at the University of Edinburgh, will facilitate collaboration on avian disease resistance, genome analysis and genome preservation. Cobb is putting almost $1 million into avian research programs at The Roslin Institute to identify innovative ways to improve avian health as well as developing unique technologies to understand and preserve the current and heritage poultry genomes. The investment creates a strategic partnership between Cobb and The Roslin Institute which leverages each world class entity s strengths. The joint partnership with Cobb is an excellent example of the kind of industrial interactions that allow The Roslin Institute s research to drive sustainable improvements in animal health and livestock productivity. One of the applications of this joint partnership is an effort to develop new technology enabling pedigree or heritage lines to be maintained without the need to physically maintain the bird stock. In addition, several projects will investigate DNA markers in the genome, targeting some of the more difficult to select for traits such as avian immunity, disease tolerance and disease resistance. The collaboration will support research by graduate students and is for an initial three years, with potential for renewal. The agreement with The Roslin Institute, which receives strategic funding from the Biotechnology and Biological Sciences Research Council, is one of more than 30 research projects that Cobb has been supporting in 18 different universities across the globe over the past five years. New partnership with Genus plc announced Efforts to improve the sustainability of livestock production will be boosted by a new research partnership. The agreement between the University of Edinburgh and Genus - an industry leader in applying genetics expertise to livestock production will look at how genetics can improve the health and welfare of production animals. This approach could include identifying genes that provide resistance to disease, to improve the quality of bred livestock and help ensure sustainable food production for a growing global and increasingly urban population. The partnership is a strategic investment and provides substantial funding for three years to support both existing and future collaborative research projects between Genus and The Roslin Institute and The Royal (Dick) School of Veterinary Studies at the University of Edinburgh. It will also fund educational initiatives. The new partnership underpins the Institute s and vet school s world-leading position in the field of livestock improvement and is central to our strategy of engaging with industry to ensure the maximum impact from our research. It provides a collaborative platform to translate insights from basic research on genetics and genomics directly into the improvement of the health and welfare of production animals. Research undertaken within the partnership will work towards the global objective of improving food security at a time when it has been acknowledged that there must be a sustainable improvement in livestock productivity.

6 Genetic chip will help salmon farmers breed better fish Atlantic salmon production could be boosted by a new technology that will help select the best fish for breeding. The development will enable salmon breeders to improve the quality of their stock and its resistance to disease. A chip loaded with hundreds of thousands of pieces of DNA each holding a fragment of the salmon s genetic code will allow breeders to detect fish with the best genes. It does so by detecting variations in the genetic code of each individual fish known as single nucleotide polymorphisms (SNPs). These variations make it possible to identify genes that are linked to desirable physical traits, such as growth or resistance to problematic diseases, for example sea lice infestations. Salmon breeders will be able to carry out the test by taking a small sample of fin tissue. The chip carries over twenty times more genetic information than existing tools. Similar chips have already transformed breeding programmes for land-farmed livestock including cattle and pigs. Salmon farming contributes around half a billion pounds to the UK economy each year and provides healthy, high quality food. Worldwide, approximately 1.5 million tonnes of Atlantic salmon are produced every year. Scientists from the University of Edinburgh s Roslin Institute and Edinburgh Genomics initiative developed the chip with researchers from the Universities of Stirling and Glasgow. They worked with industrial partners Affymetrix UK and Landcatch Natural Selection. The work was funded by the UK s innovation agency the Technology Strategy Board and the Biotechnology and Biological Sciences Research Council. Dr Ross Houston, of The Roslin Institute, said: Selective breeding programmes have been used to improve salmon stocks since the 1970s. This new technology will allow the best breeding fish to be selected more efficiently and accurately, particularly those with characteristics that are difficult to measure such as resistance to disease Dr Alan Tinch, director of genetics at Landcatch Natural Selection, said: This development takes selective breeding programmes to a whole new level. It is an extension to the selective breeding of salmon allowing more accurate identification of the best fish to create healthier and more robust offspring. Roslin scientists help decode the pig genome Researchers at The Roslin Institute headed up an international team of researchers that unravelled the genetic code of the domestic pig and the wild boars from which they are descended. The research showed that the 1 million year separation between European and Asia wild boar has resulted in significant differences in their genetic make-up. Echoes of these differences can still be seen in the differences between current day Western and Chinese breeds of domestic pigs and confirms the theory that pigs were independently domesticated in western Eurasia and East Asia. The scientists identified about 21,000 genes in the pig genome and compared these genes to their counterparts in human, mice, dogs, horses and cows. These comparisons revealed, amongst other details, that the immune response genes associated with fighting infection are evolving rapidly. The improved understanding of how these immune response genes have evolved could help combat disease and improve pig health. The research was conducted under the auspices of the Swine Genome Sequencing Consortium and led by scientists at The Roslin Institute at the University of Edinburgh, the Wellcome Trust Sanger Institute in the United Kingdom, Wageningen University in the Netherlands and the University of Illinois in the United States. The study involved more than 40 institutions in 12 countries and was funded by the United States Department of Agriculture, the European Commission, and the Biotechnology and Biological Sciences Research Council and The Wellcome Trust as well as by pig industry groups in Europe and the United States.

7 Breeding resistance to Bovine TB into cows Professor Liz Glass of The Roslin Institute has been leading a project to investigate new approaches to managing Bovine TB - a disease which cost the UK economy an estimated 90M in 2010 and which is on the rise. Professor Glass's team, which includes colleagues from Queen's University Belfast and the Agri Food and Biosciences Institute in Northern Ireland, has found that some degree of resistance to Bovine TB is inherited and the team has also identified genetic markers associated with resistance. These results mean that it might be possible to selectively breed cows which are more resistant to the disease. The group is now working with an industrial partner, DairyCo, to explore the possibility of implementing selection for increased resistance in commercial dairy cattle. The Roslin Institute Funding The Roslin Institute receives its core operational budget from the BBSRC and from the University of Edinburgh. This core support is substantially backed by individual Response Mode BBSRC funding, Wellcome Trust, Medical Research Council, Defra, Department of Health, Scottish Funding Council and the European Community. The Roslin Institute receives additional research funding from: Food Standards Agency Cystic Fibrosis Trust Meat & Livestock Commission Milk Development Council SERD DTI NIH (National Institute of Health) USA British Council Royal Society About The EBRC The Roslin Institute is a member of the Easter Bush Research Consortium (EBRC), which brings together scientists from The Roslin Institute, the Royal (Dick) School of Veterinary Studies, the Animal Sciences Researchers of Scotland s Rural College and the Moredun Research Institute. The partnerships within the EBRC, and integration with clinical practice and education in the Royal (Dick) School of Veterinary Studies, provide major opportunities for application and exploitation of the research of the consortium partners. Together, the EBRC forms one of the largest groups focussed on the biology of companion and production animals in the world. It undertakes basic and translational science to tackle pressing issues in animal genetics and genomics, development, health and welfare and their implications for human health. EBRC research has a large focus on animal and human health and this includes identifying new and emerging diseases that can pass from livestock and wild animals to humans and understanding the ways in which these diseases work. EBRC scientists with common and complimentary interests in research work closely together to develop effective disease controls and treatments, improved food safety, improved animal welfare and sustainable management of farm animals. The EBRC offers mechanisms and forums for sharing resources and expertise as well as providing an outstanding environment for teaching and training scientists at all stages of their careers. The EBRC has already established a partnership with Zoetis (formerly Pfizer Animal Health) worth at least 1.75M in research funding over 5 years, has a partnership with Genus worth 0.9M in research funding over 3 years and is negotiating similar joint R&D partnerships with several other major companies in the livestock sector (e.g. The Roslin Institute recently signed an agreement with Cobb Vantress worth almost $1 million. Easter Bush Campus Development The Roslin Institute is part of a UK Research and Innovation Campus at Easter Bush: Delivering Excellence in Animal Science and Food Security

8 The Strategy for the Growth of Easter Bush Campus The proposed Campus developments will deliver for the UK a truly outstanding and world-leading major activity in livestock research and its commercialisation. The Vision Livestock farming remains a major industry in the UK, crucial to the life and sustainability of rural communities. Sustainable increases in the productivity of the agriculture sector are required to address the future food security of UK and the world. The consolidation of The Roslin Institute, Royal Dick Vet School for Veterinary Studies (R(D)SVS) and SRUC at Easter Bush in close proximity to the Moredun Research Institute forms the Easter Bush Research Consortium (EBRC); a unique concentration of world leading in animal sciences research. The Easter Bush Campus already has the largest concentration of animal science related expertise anywhere in Europe: it aims to become internationally recognised in animal science by 2025, leading efforts to address livestock improvement, food security and alleviation of poverty in developing countries. The Easter Bush Campus aims to provide a dynamic and vibrant working environment with world class facilities equipped with the latest technologies. This allows us to continue to attract scientists and clinicians of the highest quality for our research, in order to foster and enhance our collaborative research environment which sustains and expands our links with industry. Purpose of the Easter Bush Campus To promote internationally-leading animal sciences research To facilitate interdisciplinary research and enhance our collaborative opportunities with our industrial partners. To reduce the cost of research by efficient use of shared infrastructure To generate internal seed funds for shared projects To promote effective cooperative research with other providers in the sector, notably BBSRC-funded National Institutes of Bioscience and UK Vet Schools. To attract research funding from new sources, notably European, major charity and international development agencies. To attract industry collaborators to a bio-incubator complex and to generate both joint venture and spin-out opportunities.

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