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Population Growth and Regulation Ecology Study of interrelationships between living things and their nonliving environment. Chapter 26 Ecosystems include everything, biotic and abiotic, within a given area. Biotic things are living things. Abiotic things are nonliving: soil, water, weather, etc. The interacting populations of organisms within an ecosystem form a community. Population Size Populations are all the organisms of a particular species that live within an ecosystem. The size of a population can fluctuate in response to environmental changes. The size is regulated by births, deaths, and migration in or out. Immigration: migration in to the ecosystem or population Emigration: migration out of the population Changes in Population Size = (Births - Deaths) + (Immigrants - Emigrants) A stable population has the same number of individuals joining and leaving. In most populations, immigrants and emigrants are negligible.

Changes in Population Size The birth and death rates are limited by two opposing forces within an ecosystem: Biotic potential is the maximum rate at which a population could increase. In ideal conditions, the biotic potential allows the maximum birth rate and minimum death rate. Environmental resistance limits the population s ultimate size by increasing deaths and decreasing births. Competition, predation, parasitism, and natural events are tools of environmental resistance. Population Growth The growth rate of a population is a function of the birth rate and death rate. Birth rate is the number of births per individual during a specific length of time. Death rate is the number of deaths per individual during a specific length of time. Growth Rate = Birth Rate - Death Rate R = B - D Population Growth Growth Rate = Birth Rate - Death Rate R = B - D If B is greater than D the population size increases. If D is greater than B the population will decline. Exponential Population Growth Under optimal conditions, with no environmental resistance, populations will exhibit exponential growth. In a given time period, each member of the population will produce an offspring. The population size doubles at fixed intervals, the doubling time. For example: 2 cells produce 4, 4 become 8, 8 become 16, 16 become 32, 32 become 64, etc. Most bacteria exhibit exponential growth in laboratory settings.

12000 Growth of E. coli 100000 Growth of E. coli Time (min) # Cells 0 10 30 20 60 40 90 80 120 160 150 320 180 640 210 1280 240 2560 270 5120 300 10240 Number of Cells 10000 8000 6000 4000 2000 0 0 100 200 300 400 Time (min) The typical "J curve" showing exponential growth. Time (min) # Cells 0 10 30 20 60 40 90 80 120 160 150 320 180 640 210 1280 240 2560 270 5120 300 10240 10000 1000 100 When plotted with an logarithmic y-axis, exponential growth plots as a straight line. Number of Cells 10 1 0 100 200 300 400 Time (min) Several factors affect the exponential growth curve: The age at which the organism first reproduces The frequency of reproduction The average number of offspring per reproduction The length of an organism s reproductive life span The death rate of individuals in the population The age at which the organism first reproduces affects population dynamics.

The death rate of individuals in the population affects population size. In nature, exponential growth only occurs under special circumstances and only for a limited time. Boom-and-Bust cycles occur, where populations undergo rapid growth when nutrients are available and rapid die-off when they re used up. Boom-and-Bust Cycles When specific conditions occur (temperature, moisture, nutrient availability, etc.), populations can thrive for short periods of time. Nutrients depleted Boom-and-Bust Cycles Favorable growth conditions Karenia brevis: one cause of red tides.

Predator-Prey Population Dynamics Populations of snowshoe hares and their lynx predators exhibit boom-and-bust cycles. Exponential growth can occur when environmental resistance is removed. Removal of a predator Introduction into a new ecosystem Invasive species: organisms with a high biotic potential that are introduced into ecosystems where they encounter little environmental resistance. In 1940, the whooping crane population had dropped to 20 individuals and was declared protected, removing human predators. The cane toad was introduced to Australia in 1935 to control cane beetle populations. With no predators, the cane toad population has increased exponentially.

Stabilizing Populations Most populations experience logistic population growth. Exponential growth up to the maximum number their environment can sustain, then they stabilize. The carrying capacity (K) is the maximum population size that can be sustained for an extended period of time without damage to the ecosystem. Stabilizing Populations When a population exceeds the carrying capacity of the ecosystem, the population usually crashes. Exceeding Carrying Capacity Limiting Population Size Environmental resistance ultimately limits the maximum population size. Density independent factors limit population size regardless of density. Density is the number of individuals per given area. Density dependent factors are more effective means of limiting population size.

Density-Dependent Factors Predators, organisms that feed on other organisms usually by hunting. Prey are those organisms being hunted. Predators tend to keep their prey at well below the carrying capacity of the ecosystem. Removing or introducing a new predator can alter an entire community. Density-Dependent Factors Parasites, organisms that live on larger organisms feeding off of its host without killing it. The host is the organism targeted by the parasite. Parasites spread more rapidly among dense populations. Competition for Resources Competition, interaction among individuals who attempt to use the same limited resource. Interspecific competition, among individuals of different species. Intraspecific competition, among individuals of the same species. Contest competition, use of social or chemical interactions to limit access to important resources, like marking and defending territory. How is the Human Population Changing?

How is the Human Population Changing? We discovered fire, invented tools, and built shelters allowing us to live anywhere. How is the Human Population Changing? Provided a larger and more dependable food supply. How is the Human Population Changing? Reduced disease allowing us to live longer. In developing countries, medical advances increased life span while maintaining high birth rates. Population Growth is Unevenly Distributed

Demographic Transitions in Population Growth of Nations Population Growth is Unevenly Distributed Mexico's population is growing rapidly. Population Growth is Unevenly Distributed Sweden s population has stabilized. Population Growth is Unevenly Distributed Italy s population, along with most of Europe, is shrinking. Sweden's population has reached replacement level fertility: the number of births equals the number of deaths.

United States Population Growth The U.S. is growing at a rate of 1% per year, more than any other developed country! Are We Exceeding Earth s Capacity? Amount of space and resources we need for all 6.5 billion of us is already 25% over the earth s limits. This does not taking account any space for other species! 850 million people are chronically undernourished. 1.1 billion people lack clean water. 2.6 billion have no sanitation. 2 / 3 of the world s agricultural land is suffering from erosion. Easter Island Island off the coast of Chile Geologic pollen records suggest a dense covering of forest. Humans populated the island for over 2000 years. Now, the island has no trees, and no knowledge of the ancient civilization. How did these humans exceed the carrying capacity of their island? Ecological Footprints An ecological footprint is a measure of a person s environmental impact. Energy Usage Type of home you live in Type of food you eat www.myfootprint.org US average = 609 acres each Global average = 58 acres each Earth s carrying capacity = 43 acres per person

Homework Thinking Through the Concepts, Review Question #9. Applying the Concepts #5 Calculate your ecological footprint at www.myfootprint.org. Name one thing (from the questions they ask) that you could realistically do to decrease your ecological footprint.