Developments in the Genetic Improvement of a Large Commercial Population in the New Zealand Sheep Industry

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Developments in the Genetic Improvement of a Large Commercial Population in the New Zealand Sheep Industry G B Nicoll Landcorp Farming Ltd Rotorua, New Zealand

Landcorp Farming Ltd State Owned Enterprise. Animal production business profit from commercial livestock products. 107 farms (368,000 ha). 612,000 sheep. 106,000 beef cattle. 124,000 deer. 19,000 dairy cattle.

Landcorp Farming Ltd... Sheep meat = 28% of Landcorp s operating revenue of NZ$104.3m. 385,000 slaughter lambs per year (1.5% of NZ lamb production). 412,000 breeding ewes. 1,300 new replacement sires per year. Landcorp breeds its own sires.

Corporate Sire Breeding Programmes... Financially-based breeding objective: Company objective of on-farm profitability is complementary to breeding objective. Single ownership of all livestock: Breeding programmes designed from perspective of the total commercial population. Value of breeding programmes to Landcorp = the value of revenue-earning progeny.

Corporate Sire Breeding Programmes... Large base female population: Develop new (enhance existing) breeding programmes to suit new market requirements. Use specialist resources low per head costs. Optimise animal breeding structure: Selection intensity, selection accuracy, generation interval, dissemination of genetic improvement.

Landcorp Sheep Breeding Programmes... Developments: Structure of programmes. Terminal sire programmes. - Large base population. - Rates of genetic improvement. - Use of specialist technology. - R & D.

Structure of Landcorp s Sheep Breeding Programmes

Maternal Sire Programme Terminal Sire Programme Genetic improvement Selection Males Males Females Mating system Male progeny All progeny Straightbred $ Crossbred $ $ $ $

Maternal Terminal Sheep Romney Lamb Supreme* Landmark* Texel Carwell* Beef cattle Angus Simmental Deer Red* Wapiti* * Interbred composites

Landcorp Terminal Sire Programmes

Landcorp Terminal Sire Programmes... Full birth details Weaning weight [Dam NLW, MAB] Liveweight (at ~6-7 months) U/sound muscle depth, width, fat depth CT weights of meat and fat in carcass CT eye muscle area BLUP-estimated BV s Lean Growth Index

Use of large base population...

Use of large base population... Landcorp Lamb Supreme Composite terminal sire line. Screened 501,000 ewe hoggets over 2 years. Selected top 0.8% on liveweight. Used wide range of sire breeds (3 years). Crossbred progeny = Landcorp Lamb Supreme. Continues to be non breed-specific.

Rates of genetic improvement... Landcorp Lamb Supreme Landcorp Texel

Rates of genetic improvement... Accumulated genetic gain in Landcorp terminal sire programmes (last 10 years) Landcorp Lamb Supreme Landcorp Texel Weight of lean + 2.1 kg + 2.3 kg Weight of fat + 0.01 kg + 0.06 kg L. dorsi area + 1.7 cm 2 + 1.7 cm 2 Liveweight at 8 months + 4.7 kg + 5.3 kg

Genetic Trend - Landcorp Lamb Supreme -/04 Lean and Fat Weights 2.5 2.0 BV Lean/BV Fat (kg) 1.5 1.0 0.5 0.0 Lean = +0.21 kg/yr Fat = +0.001 kg/yr -0.5 1995 1996 1997 1998 1999 2000 2001 2002 2003 2004 Year of Birth

Landcorp Texel - Genetic trend -/04 Lean and Fat Weights 2.5 2.0 BV Lean/BV Fat (kg) 1.5 1.0 0.5 0.0 Lean = +0.23 kg/yr Fat = +0.006 kg/yr -0.5 1995 1996 1997 1998 1999 2000 2001 2002 2003 2004 Year of birth

Use of specialist technology... (eg, INNERVISION TM ) Spread costs over large base population Measurement accuracy Appropriate carcass trait BVs Increased rate of genetic improvement

Estimated benefit from including CT measurements Landcorp Lamb Supreme... Lean Index ( ) 1000.0 800.0 600.0 400.0 200.0 0.0 U/S+CT: trend = +169.5 /yr U/S: trend = +100.7 /yr +316.8-200.0 1991 1992 1993 1994 1995 1996 1997 1998 1999 Year of Birth (Nicoll et al., 2002)

Specialist technology - further developments... INNERVALUE TM Kvame et al. (2004): 6 CT images to predict weight and composition of primal lamb cuts. Selection index of merit for primal cuts. Higher net benefits compared with standard 4 CT images.

Specialist technology - further developments... CT scanning vs progeny testing Discounted net benefits from two-stage selection (CT and progeny testing with 10 or 20 progeny) (Jopson et al., 2004)

Specialist technology - further developments... CT scanning young Wapiti bulls

Specialist technology - further developments... Landcorp Carwell Inseminated >300 ewes from Carwell stud, NSW, Australia.. In 4 years Identified the REM locus. Developed a marker test for the Carwell gene. Evaluated phenotypic effects of the gene. Determined its mode of inheritance.

Landcorp Carwell Effects of the Carwell gene... +10-11% L. dorsi area. +7% L. dorsi weight. Dominant inheritance. No adverse effects on L. dorsi tenderness. Frequency of Carwell allele has increased from 25% to 85% in 10 years. Parallel introgression of putative Myostatin gene variant into the Landcorp Carwell flock. (= a unique resource?)

Participation in industry initiatives... Landcorp routinely contributes data for a range of industry investigations: Sire effects on lamb survival. Sire effects on ewe litter size. Validation of markers for disease resistance (FE). Search for/fine mapping of muscling QTL. Across-flock sire evaluation (NZ Texel Society, ACE). Industry-based central progeny tests.

Summary... Landcorp Farming Ltd is a unique, large-scale commercial livestock production business. Sire breeding programmes are designed to relate directly to the company s commercial profitability. Scale advantages cost-effectively exploit application of animal breeding principles and specialist technologies.

Summary... High rates of genetic improvement in Landcorp s ram breeding programmes is only a means to an end. Profitable commercial sheep production is the measure of the value of the breeding programmes to the company.