Flight 93 National Memorial Reforestation Project (Phases I-IV)

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1 Photo credit: National Park Service Flight 93 National Memorial Reforestation Project (Phases I-IV) M.C. Tyree 1, J. Larkin 1, S. Eggerud 2, P. Angel 2, M. French 3, C. Barton 4 1 Indiana University of Pennsylvania, Department of Biology 2 Office of Surface Mining Reclamation and Enforcement 3 The American Chestnut Foundation 4 Department of Forestry, UK

2 Field collection Photo: Ian Forte (left) and Cassandra Krul (right) collecting field data at Flight 93 National Memorial. Posted at

3 Funded by: Green Forests Work The National Park Foundation (Permit #: FLNI-2015-SCI-0003) Indiana University of Pennsylvania, Department of Biology Additional thanks Leroy Renninger and MaryEllen Snyder from the NPS. Thanks to Emily Newton, Virginia Tech, The PINEMAP Undergraduate Fellowship Program for her assistance with data collection throughout the summer. Wesley Palmer and Wilson Hood for their GIS assistance throughout this project. Finally, to the hundreds of individuals that volunteered their time to help plant the site.

4 Flight 93 National Memorial Near Shanksville, PA (40.058, )

5 Flight 93 National Memorial September 10, 2011 The site was officially dedicated and opened to the public Photo credit: National Park Foundation. Accessed on at

6 Flight 93 National Memorial 1950 s until mid-1990 s Surface mined and then most of the 890 ha of was re-contoured and seeded.

7 Flight 93 National Memorial today National Park Service & Office of Surface Mining Reclamation and Enforcement teamed with others to begin reforesting sections using native woody trees and shrubs Photo credit: National Park Service

8 Reforestation Project ( ; Phases I-IV) Phase Year Area ha (ac) No. of trees I (19) 14,369 II (23) 17,300 III (28) 20,550 IV (28) 22,000 Total 39.3 (98) 74,219 > 30 spp. of native trees and shrubs Photo credit: National Park Service

9 Planting List Phase I Phase II Phase III Phase IV Total (2012) (2013) (2014) (2015) (All) Plant name # plants #/ha # plants #/ha # plants #/ha # plants #/ha # plants white pine 4, , , , ,550 red pine ,300 pitch pine ,600 Virginia pine eastern hemlock 0 0 1, ,700 red spruce ,500 Conifer tree species subtotal 5, , , , ,000 red oak 1, , , , ,600 white oak , ,100 black oak sugar maple ,070 red maple ,300 black cherry , ,900 black locust ,200 quaking aspen ,800 black walnut ,100 blackgum yellow-poplar hickory species , ,600 American elm American chestnut backcrosses * , , ,549 Deciduous tree species subtotal 6, , , , ,119 American hazelnut flowering dogwood gray dogwood silky dogwood red osier dogwood ,200 sweet American crabapple Washington hawthorn elderberry staghorn sumac mountain ash ninebark scrub oak Wildlife trees & shrubs subtotal 2, , , , ,100 GRAND TOTAL 14,369 1,869 17,300 1,895 20,550 1,824 22,000 1,968 74,219 Table. Absolute (total number) and relative (plants per hectare) abundance of woody trees and shrubs planted across the Phases I-IV of The Flight 93 National Monument Reforestation Project. Planting for each phase took place in the second half of April of each year.

10 Reforestation Project ( ; Phases I-IV) Objectives 1. Survival 2. Deer Browse 3. Competing Vegetation 4. Planting Position Summer 2015 each Phase was evaluated independently and represented 3, 2, 1, and 0.25 years following planting of Phase I, II, III, and IV, respectively.

11 Reforestation Project (2012; Phase I) Size = 7.7 ha (19 ac) No. of plots = 28 Percent sampling = 15%

12 Reforestation Project (2013; Phase II) Size = 9.1 ha (23 ac) No. of plots = 30 Percent sampling = 13%

13 Reforestation Project (2014; Phase III) Size = 11.3 ha (28 ac) No. of plots = 30 Percent sampling = 11%

14 Reforestation Project (2015; Phase IV) Size = 11.2 ha (28 ac) No. of plots = 30 Percent sampling = 11%

15 Plot Establishment Step 1: Plot boundary captured using Garmin 64s handheld GPS Step 2: Random plot centers generated using ArcGIS 10.2 Constraints > 30 meters between plot centers > 5 meters from boundary edge Phase Yr. planted Adj. area (ha) No. of plots % sample I % II % III % IV % Total %

16 Plot Establishment Step 1: Navigate to pre-established plot centers using Trimble Juno T41 GPS unit. Step 2: Drive 30 cm piece of rebar at plot center for relocating in future. Step 3: Plot photo taken Plot layout with 0.04 ha (0.10 ac) survey plot with four 1 m 2 nested subplots for percent cover determination. Step 4: Measure whole-plot data

17 Whole-Plot Measurements All planted woody plants within plot boundary were measured Measurements (Obj.) Height (Obj. 1) Basal diameter (Obj. 1) Species (Obj. 1) Vigor index (Obj. 1) Browse index (Obj. 2) Planting position (Obj. 4) Image: Posted at

18 Nested Sub-Plot Measurements Competing vegetation (Obj. 3) Categories (nearest 5%) Bare soil Rock Woody debris Grass/sedge Fern Forb Rubus spp. Woody vegetation Other Totaled 100%

19 Data Analysis Phases were treated as a separate experiments Plot-level variables that were calculated for each species (Table) Variable Description Symbol Units Species Abundance Number of individual plants for each species found within the n # or no. phase Relative abundance Number of plants relativized by plot size (0.04 ha) and expressed plants/ha #/ha (observed) on an area basis Mean height Mean height of all plants for each species within the plot Avg HT cm Minimum height Shortest individual for each species within the plot Min cm Maximum height Tallest individual for each species within the plot Max cm Mean diameter Mean basal diameter of al plants for each species within the plot Avg DIA mm Mean vigor Mean vigor value for each species within the plot. Vigor index Percent BO Percentage of plants that show no sign of deer browse % B0 % Percent B1 Percentage of plants with light deer browse % B1 % Percent B2 Percentage of plants with moderate deer browse % B2 % Percent B3 Percentage of plants with heavy deer browse % B3 % Phase-level data was averaged and standard error calculated. Treatments (position) were tested using ANOVA PDIFF Option for mean separation.

20 Overall Survival A total of 74,069 trees were planted across phases I-IV Overall survival was 55% Range was 93% - 37% within individual phases Phase I (2012) 93% (adj. 75%) Phase II (2013) 54% Phase III (2014) 41% Phase IV (2015) 37%

21 Phase I Survival (Obj. 1) + + +?? - [3 yrs.] PHASE I: Survival Planted Observed ± se Diff. Survival Distrib. Tot. Plants ± se Plant name (#/ha) (#/ha) (+/-) (%) (# plots) (#/phase) white pine ± ,920 ± 15.5 red pine ± ± 1.1 pitch pine ± ± 2.2 Virginia pine eastern hemlock red spruce Conifer tree subtotal ,403 red oak ± ,006 ± 5.9 white oak ± ± 2.6 black oak ± ± 2.3 sugar maple ± ± 2.6 red maple ± ± 11.1 black cherry ± ± 3.0 black locust ± ± 3.2 quaking aspen ± ± 2.4 black walnut blackgum ± ± 5.5 yellow-poplar hickory species American elm American chestnut backcross ± ± 2.0 Deciduous tree subtotal ,879 93% (adj. 75%) *

22 Phase II Survival (Obj. 1) [2 yrs.] PHASE II: Survival Planted Observed ± se Diff. Survival Distrib. Tot. Plants ± se Plant name (#/ha) (#/ha) (+/-) (%) (# plots) (#/phase) white pine ± ± 18.7 red pine 22 3 ± ± 0.5 pitch pine ± ± 4.1 Virginia pine eastern hemlock ± ± 4.9 red spruce Conifer tree subtotal ,900 red oak ± ± 3.9 white oak ± ± 1.5 black oak 0 1 ± ± 0.3 sugar maple ± ± 3.4 red maple ± ± 1.4 black cherry ± ± 3.3 black locust ± ± 3.0 quaking aspen ± ± 2.4 black walnut ± ± 2.7 blackgum yellow-poplar 22 7 ± ± 1.1 hickory species American elm American chestnut backcross ± ± 2.2 Deciduous tree subtotal ,789 54% *

23 Phase III Survival (Obj. 1) [1 yrs.] PHASE III: Survival Planted Observed ± se Diff. Survival Distrib. Tot. Plants ± se Plant name (#/ha) (#/ha) (+/-) (%) (# plots) (#/phase) white pine ± ± 19.8 red pine 18 2 ± ± 0.5 pitch pine ± ± 3.6 Virginia pine eastern hemlock ± ± 2.7 red spruce 71 8 ± ± 2.7 Conifer tree subtotal ,729 red oak ± ± 5.8 white oak 71 7 ± ± 1.1 black oak 0 4 ± ± 0.9 sugar maple ± ± 2.2 red maple 18 8 ± ± 1.1 black cherry ± ± 3.3 black locust ± ± 2.8 quaking aspen ± ± 2.8 black walnut 36 1 ± ± 0.4 blackgum yellow-poplar 18 6 ± ± 1.0 hickory species ± ± 1.8 American elm American chestnut backcross ± ± 4.2 Deciduous tree subtotal ,172 41% *

24 Phase IV Survival (Obj. 1) * [3 mo.] PHASE IV: Survival Planted Observed ± se Diff. Survival Distrib. Tot. Plants ± se Plant name (#/ha) (#/ha) (+/-) (%) (# plots) (#/phase) white pine ± ± 13.1 red pine pitch pine 0 1 ± * 1 9 ± 0.4 Virginia pine ± ± 2.8 eastern hemlock ± ± 2.6 red spruce ± ± 9.7 Conifer tree subtotal ,922 red oak ± ± 5.6 white oak ± ± 3.7 black oak sugar maple ± ± 2.2 red maple 18 8 ± ± 1.1 black cherry ± ± 3.3 black locust ± ± 2.7 quaking aspen ± ± 10.9 black walnut ± ± 3.4 blackgum yellow-poplar 18 6 ± ± 1.4 hickory species 98 3 ± ± 0.8 American elm American chestnut backcross ± ± 4.3 Deciduous tree subtotal ,885 37% *

25 Deer Browse (Obj. 2) Deer browse was extremely low across all four phases (3%) Phase I showed the greatest with 5% Quaking aspen (19%) and blackgum (18%) showed highest level. ***Caution, Caution, Caution, Caution, Caution, Caution, Caution***

26 Competing Vegetation (Obj. 3) Percent Cover (%) Bare Ground Years Since Planting

27 Competing Vegetation (Obj. 3) Percent Cover (%) Bare Ground Grass/Sedge Years Since Planting

28 Competing Vegetation (Obj. 3) Approx. 10% woody vegetation in year 3. Percent Cover (%) Bare Ground Forbs Grass/Sedge Years Since Planting

29 Planting Position (Obj. 4) Conifers Average plant height for bottom (B), middle (M), and top (T) planting locations for each plant group. Different letters indicate significant (P<0.05) location differences within each plant group.

30 Planting Position (Obj. 4) Across all four phases Average vigor by planting location. Different letters indicate significant (P<0.05) location differences.

31 Planting Position (Obj. 4) *Different letters indicate significant (P<0.05)

32 Future Work Summer 2017 we will remeasure phases I-IV and establish permanent plots in Phases V & VI. More intensive sampling in specially designated sections where red spruce and resistant American elm were planted (line-intersect method with multiple transects) Long-term - Continue to track stand development across all planted sites.

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